B.S. - Indiana University of Pennsylvania, Biology 1992, Minors: Anthropology and Chemistry

M.S. - Virginia Polytechnic Institute and State University, Fisheries and Wildlife Sciences, 1995

Thesis title:  Fine sediment effects on brook trout egg and alevin survival in Virginia.

Ph.D. - The Pennsylvania State University, Wildlife and Fisheries Science, 2000

Dissertation title:  Fish distribution patterns in Pennsylvania: anthropogenic influences and the status of rare fishes.